The Myriapoda (Millipedes, Centipedes) Featuring the North American Fauna     
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Bollmaniulini. Bollmaniulus sp. North America, USA, Oregon, Benton Co. Photo courtesy of W. P. Leonard.

 

Bollmaniulini. Bollmaniulus sp. North America, USA, Washington, Thurston Co., close up of two individuals in aggregation.

 

Bollmaniulini. Karteroiulus alaskanus (Cook, 1904) North America, USA, Alaska, Baranof I., Sitka. Photo courtesy of M. Goff, Sheldon Jackson College.

 

Bollmaniulini. Karteroiulus alaskanus (Cook, 1904) North America, USA, Alaska, Baranof I., Sitka. Photo courtesy of M. Goff, Sheldon Jackson College.

Bollmaniulini. North America, USA, Alaska, Prince of Wales I., Litiulus alaskanus (Cook), early instars. Photo by K. Ovaska, Aug. 2007.

 

Bollmaniulini. North America, USA, Alaska, Prince of Wales I., Litiulus alaskanus (Cook), egg chamber. Photo by K. Ovaska, Aug. 2007.

Aniulini. Aniulus (Hakiulus) d. diversifrons (Wood, 1865), gonopods of male, lateral view. North America, USA, Wisconsin, Polk Co., Osceola. Photo courtesy of J. Wheeler.

Aniulini. Aniulus (Hakiulus) d. diversifrons (Wood, 1865), enlarged, clasper-like 1st legs of male. North America, USA, Wisconsin, Polk Co., Osceola. Photo courtesy of J. Wheeler.

Aniulini. Oriulus venustus (Wood, 1864). North America, USA, Wyoming, Yellowstone National Park near Yellowstone River. Photo by S. Hardy; hand of M. Dowling.

 

 

 

 

 

Bollmaniulini. Bollmaniulus sp. in typical resting coil seen from lateral perspective. North America, USA, Washington, Thurston Co., Capitol Forest. Photo courtesy of W.P. Leonard.

Bollmaniulini. Bollmaniulus sp. North America, USA, Washington, Thurston Co., aggregation on porch ceiling at house in Lacey. Photo courtesy of M. Kraft.

Bollmaniulini. Karteroiulus alaskanus (Cook, 1904); note that the coil forms a raised, rather than a flat, spiral. North America, USA, Alaska, Baranof I., Sitka. Photo courtesy of M. Goff, Sheldon Jackson College.

 

Aniulini. Arvechambus hummi Causey, 1963, hermaphroditic spmn., close up of genitalic segments, ventral view. North America, USA, Florida, Alachua Co., Gainesville. Photo by A. E. Bogan.

Aniulini. Aniulus ?garius(Chamb., 1910). North America, USA, Wisconsin, Polk Co. Photo courtesy of J. Wheeler.

Female parajulid with Gordius/ Horse hair worms (Nematomorpha) protruding through anus. North America, USA, Wisconsin, St. Croix R.  Photo by J. Wheeler.

Aniulini. Aniulus garius (Chamb., 1912) M. North America, USA, Wyoming, Weston Co. Photo courtesy of S.-K. Wu & Univ. Colorado Museum of Natural History

 

 

 

 

 

 

 

 

 

Representative Parajulidae, the dominant North American millipede family. Note enlarged 1st segment (or collum), with which it burrows.

 

Bollmaniulini.  Bollmaniulus sp. North America, USA, Washington, Thurston Co., Capitol State Forest. Photo courtesy of W. P. Leonard.

 

Aniulini. ?Gyniulus sp. North America, USA, Florida, Pinellas Co., St. Petersburg. Photo courtesy of G. B. Edwards.

 

Aniulini. Arvechambus hummi Causey, 1963. Anterior end of hermaphroditic spmn., lateral view. North America, USA, Florida, Alachua Co., Gainesville. Photo by A. E. Bogan.

 

Aniulini. Arvechambus hummi Causey, 1963, anterior end of hermaphroditic spmn., ventral view. North America, USA, Florida, Alachua Co., Gainesville. Photo by A. E. Bogan.

Bollmaniulini. North America, USA, Alaska, Prince of Wales I., Litiulus alaskanus (Cook). Photo by K. Ovaska, Aug. 2007.

Bollmaniulini. Anterior end of ?Bollmaniulus sp. F. North America, Canada, British Columbia, Vernon. Infesting residence. Photo by W. Strong.

Partial dissection of female parajulid infested with Nematomorpha (Gordius/Horse hair worms); anterior end removed to show coiled worm in body cavity and one protruding through anus. North America, USA, Wisconsin, St. Croix R.  Photo by J. Wheeler.

Complete dissection of Nematomorpha (Gordius/Horse hair worms) from infested female parajulid. North America, USA, Wisconsin, St. Croix R.  Photo by J. Wheeler.

Ptyoiulus impressus (Say, 1821). Anterior end of male showing greatly enlarged 1st legs. North America, USA, North Carolina, Watauga Co. Photo by D. Hennen.

 

 

 

 

 

 

 

 

Bollmaniulini. Bollmaniulus sp. North America, USA, Washington, Thurston Co., Lacey. Three individuals from aggregation. Photo courtesy of M. Kraft.

 

Aniulini. ?Gyniulus sp., anterior end. North America, USA, Florida, Pinellas Co., St. Petersburg. Photo courtesy of G. B. Edwards.

 

Aniulini. ?Gyniulus sp., caudal end. North America, USA, Florida, Pinellas Co., St. Petersburg. Photo courtesy of G. B. Edwards.

 

Aniulini. Arvechambus hummi Causey, 1963, hermaphroditic spmn., close up of genitalic segments, lateral view. North America, USA, Florida, Alachua Co., Gainesville. Photo by A. E. Bogan.

Bollmaniulini. North America, Canada, British Columbia, Prince Rupert vicinity, Litiulus alaskanus (Cook), adult female beside egg chamber. Photo by E. Havard, Aug. 2007.

Bollmaniulini. ?Bollmaniulus sp. F. North America, Canada, British Columbia, Vernon. Infesting residence. Photo by W. Strong.

Bollmaniulini. ?Bollmaniulus sp. F compared to Canadian penny. North America, Canada, British Columbia, Vernon. Infesting residence. Photo by W. Strong.

Nematomorpha (Gordius/Horse hair worms) from infested female parajulid separated; lengths 7.0 & 7.5 cm. North America, USA, Wisconsin, St. Croix R.  Photo by J. Wheeler.

Uroblaniulini. Uroblaniulus ?canadensis (Newport, 1844) F. Note decurved, "hook-like" epiproct. North America, Canada, Québec, Montreal vicinity. Photo courtesy of P.-M. Brousseau.

 

 

 

 

 

 
IntroMillipedesCentipedesPaleozoic FossilsDifferencesCongressesReferencesMyriapodologists-DeceasedMyriapodologists-LivingPhoto GalleryAlaska & Northwestern CanadaBiogeographyEvolutionary Diagrams